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Cranial osteology and phylogenetic position of the theropod dinosaurProceratosaurus bradleyi(Woodward, 1910) from the Middle Jurassic of England

2009/11/04 by Oliver W. M. Rauhut, Angela C. Milner, Scott Moore-Fay · 120 citations
Earth and Planetary Sciences · Environmental Science · #Paleontology and Evolutionary Biology #Evolution and Paleontology Studies #Ichthyology and Marine Biology #Osteology #Biology #Phylogenetic tree #Anatomy #Paleontology #Evolutionary biology #Zoology #Genetics

paper · pdf · doi:10.1111/j.1096-3642.2009.00591.x

published in Zoological Journal of the Linnean Society 158(1), 155-195 (Oxford University Press)

openalex publication_date 2009/11/04 · openalex created_date 2025/10/10 · openalex updated_date 2026/06/23

Abstract

The cranial osteology of the small theropod dinosaur Proceratosaurus from the Bathonian of Minchinhampton, England, is described in detail, based on new preparation and computed tomography (CT) scan images of the type, and only known, specimen. Proceratosaurus is an unusual theropod with markedly enlarged external nares and a cranial crest starting at the premaxillary–nasal junction. The skull is highly pneumatic, with pneumatized nasals, jugals, and maxillae, as well as a highly pneumatic braincase, featuring basisphenoid, anterior tympanic, basipterygoid, and carotid recesses. The dentition is unusual, with small premaxillary teeth and much larger lateral teeth, with a pronounced size difference of the serrations between the mesial and distal carina. The first dentary tooth is somewhat procumbent and flexed anteriorly. Phylogenetic analysis places Proceratosaurus in the Tyrannosauroidea, in a monophyletic clade Proceratosauridae, together with the Oxfordian Chinese taxon Guanlong. The Bathonian age of Proceratosaurus extends the origin of all clades of basal coelurosaurs back into the Middle Jurassic, and provides evidence for an early, Laurasia-wide, dispersal of the Tyrannosauroidea during the late Middle to Late Jurassic.

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